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Pestle Puffball (Handkea excipuliformis)

How to identify Pestle Puffball

Handkea excipuliformis

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To correctly identify Pestle Puffball (Handkea excipuliformis), check each of these features in order. Misidentification is the most common cause of mushroom poisoning, so cross-reference multiple sources.

Step-by-step identification

  1. 1

    Cap

    Shape, color, surface texture, and diameter.

    The Pestle Puffball does not have a distinct cap. The entire fruit body is a single structure.

  2. 2

    Gills

    Attachment to stem, color, and spacing.

    Absent. Spores are produced inside the gleba.

  3. 3

    Stem

    Length, shape, surface, and any ring.

    Absent or rudimentary. If present, it is a short, sterile base.

  4. 4

    Flesh

    Color, texture, and bruising reaction.

    The flesh (gleba) is white and firm when young, turning yellowish and then brownish as it matures. Only consume when the flesh is uniformly white.

  5. 5

    Fruiting body

    Overall growth pattern.

    Pear-shaped to club-shaped, 5-20 cm tall, 4-10 cm wide. White when young, becoming brownish with age.

  6. 6

    Habitat

    Where it grows and on what substrate.

    Found in meadows, fields, and open woodlands, often growing on the ground.

Key Facts

Cap surfaces
Smooth when young, may become cracked with age
Growth Form
Solitary or in groups
Spore Print
Olive-brown
Body color
White to brownish
Odor
Mild, mushroomy
Flesh bruises
Does not bruise
Height
5-20 cm
Cap Diameter
N/A (no distinct cap)

Fruit body characteristics

  • Gill attachment: Absent (Gleba)
  • Stem shape: Absent or rudimentary
  • Stem surface: N/A
  • Ring: Absent
  • Stem cross-section: N/A

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